The Small Mountain Ringlet Erebia epiphron
is a speciality of the Lake District, this being the only
place in England where it occurs. It is an enigmatic species
living on the high fells but only flying in sunshine! Its
larval foodplant, Matt Grass, is everywhere, but the
butterfly is found only in specific areas, apparently linked
to the geology, soil type and possibly aspect.
Colonies show a marked preference for the west and south-west
of the region, especially around Honister, the Langdales and Wrynose
where numbers have traditionally been highest and are holding
up well. Colonies to the east, around Ullswater, Haweswater and
Kirkstone, are fewer in number, generally smaller and more isolated.
The Red Screes colony, where the species was first discovered
in Britain in the early 1800's, now appears to be very small.
The species appears in the Lake District earlier than many books
suggest. Numbers peak very quickly, often around the last week
of June, followed by a fairly long tail. Data held at Tullie House,
Carlisle, covering the whole of the last century provide the following
analysis:
Week
beginning |
5th
June |
12th
June |
19th June |
26th
June |
3rd
July |
10th
July |
17th July |
23rd
July |
No.
of Records |
2 |
4 |
16 |
46 |
31 |
25 |
6 |
1 |
Hot
weather during the flight period may restrict its flight to no
more than a fortnight, whereas cool cloudy weather makes for a
longer tail but little opportunity for flight and observation.
Predation by birds is also heavy during sunny periods, perhaps
also contributing to a shorter tail to the flight period. The
lower altitude colonies tend to emerge a little earlier.
One
feature of the species that experienced watchers have noted is
that the peak flight period does seem to vary backwards and forwards
from year to year. This is a complex behaviour that is not simply
linked to spring temperatures. For example, early emergence has
been known following a cold wet spring and in a cold wet summer.
It might be that this is a consequence of the weather in previous
years. From September to April the half-grown caterpillars hibernate
deep in tussocks, often covered with a protective blanket of snow.
It is believed that adverse weather in spring (or perhaps even
in late summer of the previous year) may delay development so
much that the larval stage lasts two years. In that event, development
would be advanced in the second year leading to early pupation
and emergence of adults, even if the current season's weather
is unfavourable.
In
short, this is a frustrating species for the casual observer to
plan to see and why records over the years are relatively sparse,
despite the increasing popularity of fell-walking!!!
Colonies
are normally found above 550 metres, although several are well
below this and one very productive site is known at a mere 250
metres:
Altitude
(m) |
250 |
400+ |
450+ |
500+ |
550+ |
600+ |
650+ |
No.
of Colonies |
1 |
4 |
1 |
3 |
12 |
8 |
11 |
In
spite of its preference for altitude the species does avoid the
summit tops, as it is nearly always found in sheltered nooks and
crannies or flushes at the base of crags. One
recent piece of research has suggested that British colonies have
moved 150 metres (500 feet) higher on average in recent years,
but this is improbable for the Lake District as it would place
most colonies on the mountain summits, which are too exposed.
Where
the species does occur, it can be very abundant and numbers can
be very high. One researcher, having camped out at a site, estimated
a total emergence over 12 days at 9000 individuals. The casual
observer will see nothing like this quantity however. Females
fly very little and, except in periods of prolonged hot sunshine,
males can spend much of their time in the vegetation. The database
of all known records for 1990-2006 (which was used to compile
the distribution maps below) contains only three records where
more than 100 individuals were present!
Known
sites are found only on certain of the Borrowdale Volcanic rocks
- the Skiddaw and Silurian Slates have no colonies. It may be
that the areas avoided, being more acidic, are mineral deficient
(especially in calcium). Matt Grass is so lacking in mineral nutrition
that even the sheep leave it alone!
Plants
on which the Mountain Ringlet nectars, such as wild thyme, tormentil
and meadow buttercup, tend to be found at the base of crags and
gullies, where mineral enrichment occurs. They would also be absent
on the summit tops, making this another reason why the species
is unlikely to move higher in the Lakes.
There
have been no records ever from some fells (although
well walked) such as Great Gable, Bowfell and the Band, The Fairfield
Horseshoe, Skiddaw and Matterdale Common, despite the abundance
of Matt Grass throughout. The species has many strong colonies
just to the north of the Wrynose Pass, but curiously has never
crossed the road to the Coniston range! There appears to be a
very strong geological link to its habitat requirements (see map
below). If you are walking in the Lake District in June or July
please look carefully and report any sightings via this website.
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|
1990
- 2006 Mountain Ringlet colonies (red dots) and ALTITUDE.
Records of less than three are omitted. |
Mountain
Ringlet colonies and GEOLOGY (green dots) - pink and lilac
are the Borrowdale volcanic rocks, red is granite. |
N.B. AN ENLARGED DISTRIBUTION MAP CAN BE SEEN BY
CLICKING ON THE ALTITUDE MAP
|
The
colonies in the west of the Lake District could all be considered
to be one super-colony as they are all inter-connected by the
ridges that radiate out from the central dome of the Scafells
(very noticeable on the altitude map above). A little interchange
probably occurs here between "colonies", leading to
genetic mixing and stronger individuals. It may be the case that
colonies in the east will continue to decline through lack of
genetic mixing caused by their greater isolation from each other.
Within
a colony there are genetic differences of size, brightness and
number of spots. English specimens differ slightly from those
of Scotland, which differ again from European specimens. Perhaps
genetic differences between the west and east Lake District colonies
may have occurred as these have become increasingly polarised,
even if there are no visible differences?
There
has been much speculation and warnings about the potential loss
of an alpine species like the Mountain Ringlet as global warming
occurs. It is just that, speculation, as so little is known about
the habitat requirements and ecology of this species. One of the
strongest colonies in the Lakes is the one found at lowest altitude
(250m) and on the milder west coast, where it enjoys the benefits
of warm Gulf Steam air and higher humidity - just the sort of
conditions Global warming is supposed to bring. This colony has
been observed for over 60 years, with little apparent change in
strength. On the other hand, a colder, drier climate is found
on the Eastern fells and should in theory favour an alpine species,
but these colonies have declined dramatically in the last 60 years,
although I suspect this is not in fact weather related.
The
trends predicted from Global warming of warmer wetter weather
may be much less significant than other requirements for the survival
of the species. Could it be that the amount of snow cover becomes
critical? At low altitudes this protective cover is not present
and appears not to be needed. At high altitude it may be beneficial,
in which case the species may well need to move to higher ground,
as appears to be the case in Scotland but this is not possible
in the Lake District.
All
in all, the Lake District Mountain Ringlets are a fascinating
and unsolved enigma! If you have any comments on the above theories
please get in touch.
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